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These homeotic genes are ordered on the DNA in the same way that they are expressed along the body axis during development.
The discovery of the genes that govern embryonic development was made possible by a little monster: a fruit fly (Drosophila) with an additional set of wings.
The evolutionary significance of developmental limitation has long been debated. Constrained advocates have been skeptical of what they regard as neo-Darwinians' panselectionist approach.
However, such supporters have been chastised for taking an overly simplistic view of Neo-Darwinism. Some argue that neo-Darwinian theory includes developmental constraints.
Two factors have limited progress in determining the evolutionary function of developmental constraints: terminological issues and a lack of clear, unambiguous instances.
I address both of these concerns in this article. I provide definitions that correspond to the majority of prior usage, but not all.
I then discuss the example of centipede segment number constraint, which is particularly clear in that an alternative selective explanation for the observed pattern of character states is highly implausible.
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